They communicate through body language and vocalizations from "summary" of The Hidden Life of Wolves by Jim Dutcher,Jamie Dutcher,James Manfull
Wolves are masters of nonverbal communication. They convey a wealth of information through their body language and vocalizations. Their tails, ears, and posture all play a crucial role in expressing their feelings and intentions to other pack members. A wolf's body language can signal aggression, submission, playfulness, or fear. When wolves interact with one another, they use a complex system of vocalizations to convey different messages. From howls to whimpers, each sound has a specific meaning that other pack members understand intuitively. For example, a low growl may signal a warning to stay away, while a high-pitched yip can indicate excitement or joy. In addition to body language and vocalizations, wolves also use scent to communicate with one another. They mark their territory with urine and scat, leaving behind a trail of information for other wolves to decipher. By sniffing these scent...
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